Beaver
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| Beavers Fossil range: Late Miocene - Recent | ||||||||||||
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American Beaver | ||||||||||||
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| C. canadensis |
Beavers are semi-aquatic rodents native to North America and Europe. They are the only living members of the family Castoridae, which contains a single genus, Castor. Genetic research has shown the European and North American beaver populations to be distinct species and that hybridization is unlikely.
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[edit] General
Beavers are best known for their natural trait of building dams in rivers and streams, and building their homes (known as beaver lodges) in the eventual pond. They are the second-largest rodent in the world (after the capybara).
Beavers continue to grow throughout life. Adult specimens weighing over 25 kg (55 lb) are not uncommon. Females are as large as or larger than males of the same age, which is uncommon among mammals.
[edit] Species
[edit] European Beaver
The European Beaver (Castor fiber) was hunted almost to extinction in Europe, both for fur and for castoreum, a secretion of its scent gland believed to have medicinal properties. However, the beaver is now being re-introduced throughout Europe. Several thousand live on the Elbe, the Rhone and in parts of Scandinavia. A thriving community lives in north east Poland, and the European Beaver also returned to the Morava River banks in Slovakia and the Czech Republic. They have been reintroduced in Bavaria, The Netherlands and Serbia (Zasavica bog) and are spreading to new locations.
The beaver finally became extinct in Great Britain in the sixteenth century: Giraldus Cambrensis reported in 1188 (Itinerarium ii.iii) that it was to be found only in the Teifi in Wales and in one river in Scotland, though his observations are clearly first hand. In October 2005, six European beavers were re-introduced to Britain in Lower Mill Estate in Gloucestershire, and there are plans for re-introductions in Scotland and Wales.[1]
[edit] Giant beaver
The North American Giant beaver (Castoroides ohioensis) was one of largest rodents that ever evolved. It disappeared, with other large mammals in the Holocene extinction event, which began about 13,000 years ago.
[edit] Habitat
The habitat of the beaver is the riparian zone inclusive of stream bed. The habit of the beaver for hundreds of thousands of years in the Northern Hemisphere has been to keep these watery systems healthy and in good repair, although to a human observer, seeing all of the downed trees, it might sometimes seem that the beavers are doing just the opposite. The beaver works as a keystone species in an ecosystem by creating wetlands that are used by many other species. Next to humans, no other extant animal does more to shape its landscape.[citation needed] Introduced to an area without its natural predators, as in Tierra del Fuego, beavers have flooded thousands of acres of land and are considered an unstoppable plague. One notable difference in Tierra del Fuego from most of North America is that the trees found in Tierra del Fuego do not coppice as do willows, poplars, aspens, and other North American trees. Thus the "damage" by the beavers seems more severe.
[edit] Dams
Beaver dams are created both as a protection against predators, such as coyotes, wolves and bears, and to provide easy access to food during winter. Destroying a beaver dam without removing the beavers takes a lot of effort, especially if the dam is downstream of an active lodge. Beavers can rebuild such primary dams overnight, though they may not defend secondary dams as vigorously.
Beavers have been known to build very large dams.[2] The largest known was discovered near Three Forks, Montana, and was 2,140 feet long, 14 feet high, and 23 feet thick at the base.[3] When objectionable beaver flooding occurs, modern water level control devices can be installed for a cost-effective and environmentally sound solution. Unwanted damage to trees can be prevented by wrapping chicken wire or sheet metal around the base of trees.[4]
It is primarily the sound of water in motion that stimulates the beavers to build. However, studies involving beaver habitual activities have indicated that beavers may respond to an array of stimuli, not just the sound of running water. In two experiments Wilson (1971) and Richard (1967, 1980) demonstrate that, although beavers will pile material close to a loudspeaker emitting sounds of water running,[5] they only do so after a considerable period of time. Additionally the beavers, when faced with a pipe allowing water to pass through their dam, eventually stopped the flow of water by plugging the pipe with mud and sticks. The beavers were observed to do this even when the pipe extended several meters upstream and near the bottom of the stream and thus produced no sound of running water. Beavers normally repair damage to the dam and build it higher as long as the sound continues. However, in times of high water, they often allow spillways in the dam to flow freely.
Beaver dams can be disruptive; the flooding can cause extensive property damage, and when the flooding occurs next to a railroad roadbed, it can cause derailments by washing-out under the tracks, or when a beaver dam bursts and the resulting flash flood overwhelms a culvert. This disruption is not limited to human geography; beavers can destroy nesting habitat for endangered species, and often destroy mature trees for which they have no use.
On the other hand, dam building is extremely beneficial in restoring wetlands. Such wetland benefits include flood control downstream, biodiversity (by providing habitat for many rare as well as common species), and water cleansing, both by the breakdown of toxins such as pesticides and the retention of silt by beaver dams. Over the eons, this collection of silt produces the rich bottom land so sought after by farmers. Beaver dams also reduce erosion as well as decrease the turbidity that is a limiting factor for much aquatic life. While beavers can create damage, part of the problem is one of perception and time scale. Such damage as the undermining of a roadway or the drowning of some trees is very visible shortly after the beginning of beavers' activity in an area. The benefits, mentioned below, are long-term and not easily seen except by someone who is monitoring a catchment and realizes the beneficial effects of beaver dams.
[edit] Flood control
A beaver dam has a certain amount of freeboard above the water level. When heavy rains occur, the pond fills up and the dam gradually releases the extra stored water. Often this is all that is necessary to reduce the height of the flood wave moving down the river, and will reduce or eliminate damage to human structures. Flood control is achieved in other ways as well. The surface of any stream intersects the surrounding water table. By raising the stream level, the gradient of the surface of the water table above the beaver dam is reduced, and water near the beaver dam flows more slowly into the stream. This helps in reducing flood waves, and increases water flow when there is no rain. Beaver dams also smooth out water flow by increasing the area wetted by the stream. This allows more water to seep into the ground where its flow is slowed. This water eventually finds its way back to the stream. Rivers with beaver dams in their head waters have lower high water and higher low water levels.
[edit] Wetland creation
If a beaver pond becomes too shallow due to the settling of sediment, or if the tree supply is depleted, beavers will abandon the site. Eventually the dam will be breached and the water will drain out. The rich thick layer of silt, branches, and dead leaves behind the old dam is the ideal habitat for wetland species. Many of them will have been on the fringes of the pond. Wetlands have significant environmental benefits.
[edit] The grazing meadow (vega)
As the wetland fills and dries out, pasture species colonize it and it becomes a meadow suitable for grazing. In an area with nothing but forest down to the stream edge, this provides a valuable niche for many animals which otherwise would be excluded.
[edit] The riverine forest
Finally the meadow will be colonized by riverine trees, typically aspens, willows and such species which are favoured by the beaver. Beavers are then likely to recolonize the area, and the cycle begins again.
[edit] Bottom land
As related above, each time this process repeats itself another layer of rich organic soil is added to the bottom of the valley. The valley slowly fills and the flat area at the bottom gets wider. Research is sparse on this topic, but it seems likely that much of the fabled bottom land in North America was created, or at least added to, by the efforts of the generations of beavers that lived there.
[edit] Nutrient removal
The removal of nutrients from the stream flow by beaver ponds is an interesting and very valuable process. Farming along the banks of rivers often increases the loads of phosphates, nitrates and other nutrients, causing problems downstream when this water is extracted for drinking. Besides silt, the beaver dam collects twigs and branches from the beavers' activity and leaves, notably in the fall. The main component of this material is cellulose, a polymer of β-glucose monomers (This creates a more crystalline structure than is found in starch, which is composed of α-glucose monomers. Cellulose is a type of polysaccharide.) Many bacteria produce cellulase which can split off the glucose and use it for energy. Just as algae get their energy from sunlight, these bacteria get their energy from cellulose, and they form the base of a very similar food chain. However, a source of energy is not enough for growth. These bacterial populations face serious shortages of nitrous and phosporous compounds, and will absorb these nutrients as they pass by in the water stream. In this way, these and other nutrients are fixed into the beaver pond and the surrounding ecology, and are removed from the stream.
[edit] Pesticide and herbicide removal
Agriculture also introduces herbicides and pesticides into our streams. Bacteria are an extremely variable lot and some of these toxicants are metabolized and decomposed by the bacteria in the cellulose-rich bottom of a beaver dam.
[edit] Denitrification
Some scientists believe that the nitrate cascade, the production of far more fixed nitrogen than the natural cycles can turn back into nitrogen gas, may be as much of a problem to our ecology as carbon dioxide production. It is likely, but not proven, that beaver dams along a stream may contribute to denitrification (the removal of nitrogen). In sewage plants, denitrification is achieved by passing the water through successive aerobic and anaerobic stages. Under a beaver dam, as the water seeps down into the soil, the oxygen is used up by the fauna in the rich organic layer. At some point all the oxygen is used up and the soil becomes anaerobic. This water eventually finds its way into the stream and into another beaver dam. This aerobic, anaerobic cycle continues all the way down the stream and denitrification is a likely result.
[edit] Lodges
The ponds created by well-maintained dams help isolate the beavers' home, their lodge, which is also created from severed branches and mud. The lodge has underwater entrances to make entry nearly impossible for any other animal (however, muskrats have been seen living inside beaver lodges with the beavers who made it). A very small amount of the lodge is actually used as a living area. Contrary to popular belief, beavers actually dig out their den with an underwater entrance after they finish building the dam and lodge structure. There are typically two dens within the lodge, one for drying off after exiting the water, and another, drier one where the family actually lives.
[edit] Danger signal
When startled or frightened, a swimming beaver will rapidly dive while forcefully slapping the water with its broad tail. This creates a loud 'slap', audible over large distances above and below water. This noise serves as a warning to other beavers in the area. Once a beaver has made this danger signal, all nearby beavers will dive and may not reemerge for some time.
[edit] Uses
Beaver pelts were used for barter by Native Americans in the 17th century to gain European goods. They were then shipped back to Great Britain and France where they were made into clothing items. Widespread hunting and trapping of beavers led to their endangerment. Eventually, the fur trade fell apart due to declining demand in Europe and the takeover of trapping grounds to support the growing agriculture sector. A small resurgence in beaver trapping has occurred in some areas where there is an over-population of beaver; trapping is only done when the fur is of value, and normally the remainder of the animal is also utilized as animal feed.
Both beaver testicles and castoreum, a bitter-tasting secretion with a slightly fetid odor contained in dried preputial or vaginal follicles of male or female beaver, have been articles of trade for use in traditional medicine. Yupik (Eskimo) medicine used dried beaver testicles like willow bark to relieve pain.[6] Beaver testicles were exported from Levant (a region centered on Israel) from the tenth to nineteenth century.[7] Claudius Aelianus comically described beavers chewing off their testicles to preserve themselves from hunters.[8] European beavers (Castor fiber) were eventually hunted nearly to extinction in part for the production of castoreum, which as used as an analgesic, anti-inflammatory, and antipyretic. Castoreum was described in the 1911 British Pharmaceutical Codex for use in dysmenorrhea and hysterical conditions (i.e. pertaining to the womb), for raising blood pressure and increasing cardiac output. The activity of castoreum has been credited to the accumulation of salicin from willow trees in the beaver's diet, which is transformed to salicylic acid and has an action very similar to aspirin.[9] Castoreum continues to be used in perfume production.
[edit] Beavers in culture
Popular western culture typically depicts the animal positively, as a good-natured and industrious character.
- The North American Beaver (C. canadensis) is the national animal of Canada; it is depicted on the Canadian five-cent piece and was on the first Canadian postage stamp, the Three-Penny Beaver. As a national symbol, the beaver was chosen to be the mascot of 1976 Summer Olympics held in Montreal with the name "Amik" ("beaver" in Algonquin). The main reason why the beaver is the national symbol of Canada is because of the beaver fur. In the late 1600s and early 1700s, the fashion of the fur top-hats came out, which convinced the first European explorers to hunt in the deep Canadian wilderness for beavers. It is also the symbol of many units and organizations within the Canadian Forces, such as on the cap badges of the Royal 22e Régiment and the Canadian Military Engineers. However, beavers are considered a pest by some people.
- Mr. and Mrs. Beaver who are important heroic characters in the classic fantasy novel, The Lion, the Witch and the Wardrobe.
- The beaver's habits, habitat and conservation status (as of 1908) are recurring themes in The Tent Dwellers, by Albert Bigelow Paine. Lillian Hoban's Charlie the Tramp is a children's book about a young beaver and his family.
- There is typically a Beaver Patrol in the Boy Scouts of America's Wood Badge adult-leadership training program. A North American Beaver in Calgary, Alberta
- In the United States, Oregon is known as the "The Beaver State." The beaver is the state animal. It is also the mascot of Oregon State University. It is the state mammal of New York (after the historical emblem of New Netherland). It also appears on New York City's coat of arms, seal and flag because of the importance of the fur trade in initial settlement of the region.
- Due to its engineering capabilities, the beaver serves as the mascot of the Massachusetts Institute of Technology, California Institute of Technology, Oregon State University, and the University of Toronto. It is also an emblem for London School of Economics and the name of its student newspaper, The Beaver.
- In the 17th century, based on a question raised by the Bishop of Quebec, the Roman Catholic Church ruled that the beaver was a fish for purposes of dietary law. Therefore, the general prohibition on the consumption of meat on Fridays during Lent does not apply to beaver meat.[10][11][12] The legal basis for the decision probably rests with the Summa Theologica of Thomas Aquinas, which bases animal classification as much on habit as anatomy.[13]
- In the Cheese Shop sketch of Monty Python's Flying Circus, a famished customer asks the proprietor of a cheese shop for any one of forty-two different kinds of cheese, including the nonexistent Venezuelan Beaver Cheese. Venezuela has no indigenous beavers.
- Nickelodeon aired The Angry Beavers, a popular children's television show.
- In computability theory, a Busy Beaver is a Turing machine which does as much work as possible given the size of its tape alphabet and underlying state machine.
- Beaver is also a popular slang term used to describe the female genitalia.
- Bibarel from Pokemon.
[edit] 1911 encyclopedia text
The following text is taken from the 1911 Encyclopedia Brittannica.
Beaver, the largest European aquatic representative of the mammalian order RODENTIA, easily recognized by its large trowel-like, scaly tail, which is expanded in the horizontal direction.
The word is descended from the Aryan name of the animal, cf. Sanskrit babhru's, brown, the great ichneumon, Lat. fiber, Ger. Biber, Swed. bäver, Russ. bobr'; the root bhru has given "brown," and, through Romanic, "bronze" and "burnish."
The true beaver (Castor fiber) is a native of Europe and northern Asia, but it is represented in North America by a closely-allied species (C. canadensis), chiefly distinguished by the form of the nasal bones of the skull.
Beavers are nearly allied to the squirrels (Sciuridae), agreeing in certain structural peculiarities of the lower jaw and skull. In the Sciuridae the two main bones (tibia and fibula) of the lower half of the leg are quite separate, the tail is round and hairy, and the habits are arboreal and terrestrial. In the beavers or Castoridae these bones are in close contact at their lower ends, the tail is depressed, expanded and scaly, and the habits are aquatic.
Beavers have webbed hind-feet, and the claw of the second hind-toe double. They have poor eyesight, but a keen sense of hearing, smell, and touch.
In length beavers—European and American—measure about 2 ft. exclusive of the tail, which is about 10 inches long. They are covered with a fur to which they owe their chief commercial value; this consists of two kinds of hair—the one close-set, silky and of a greyish colour, the other much coarser and longer, and of a reddish brown.
Beavers are essentially aquatic in their habits, never travelling by land unless driven by necessity. Formerly common in England, the European beaver has not only been exterminated there, but likewise in most of the countries of the continent, although a few remain on the Elbe, the Rhone and in parts of Scandinavia. The American species is also greatly diminished in numbers from incessant pursuit for the sake of its valuable fur.
Beavers are sociable animals, living in streams, where, so as to render the water of sufficient depth, they build dams of mud and of the stems and boughs of trees felled by their powerful incisor teeth. In the neighbourhood they make their "lodges," which are roomy chambers, with the entrance beneath the water. The mud is plastered down by the fore-feet, and not, as often supposed, by the tail, which is employed solely as a rudder.
They are mainly nocturnal, and subsist chiefly on bark and twigs or the roots of water plants.
The dam differs in shape according to the nature of particular localities. Where the water has little motion it is almost straight; where the current is considerable it is curved, with its convexity towards the stream. The materials made use of are driftwood, green willows, birch and poplars; also mud and stones intermixed in such a manner as contributes to the strength of the dam; but there is no particular method observed, except that the work is carried on with a regular sweep, and that all the parts are made of equal strength.
"In places," writes Hearne, "which have been long frequented by beavers undisturbed, their dams, by frequent repairing, become a solid bank, capable of resisting a great force both of ice and water; and as the willow, poplar and birch generally take root and shoot up, they by degrees form a kind of regular planted hedge, which I have seen in some places so tall that birds have built their nests among the branches."
Their houses are formed of the same materials as the dams, with little order or regularity of structure, and seldom contain more than four old, and six or eight young beavers. It not unfrequently happens that some of the larger houses have one or more partitions, but these are only posts of the main building left by the builders to support the roof, for the apartments have usually no communication with each other except by water.
The beavers carry the mud and stones with their fore-paws and the timber between their teeth. They always work in the night and with great expedition. They cover their houses late every autumn with fresh mud, which, freezing when the frost sets in, becomes almost as hard as stone, so that neither wolves nor wolverines can disturb their repose.
The favourite food of the American beaver is the water-lily (Nuphar luteum), which bears a resemblance to a cabbage-stalk, and grows at the bottom of lakes and rivers. Beavers also gnaw the bark of birch, poplar and willow trees; but during the summer a more varied herbage, with the addition of berries, is consumed.
When the ice breaks up in spring they always leave their embankments, and rove about until a little before the fall of the leaf, when they return to their old habitations, and lay in their winter stock of wood. They seldom begin to repair the houses till the frost sets in, and never finish the outer coating till the cold becomes severe. When they erect a new habitation they fell the wood early in summer, but seldom begin building till towards the end of August.
Castoreum is a substance contained in two pear-shaped pouches situated near the organs of reproduction, of a bitter taste and slightly foetid odour, at one time largely employed as a medicine, but now used only in perfumery.
Fossil remains of beavers are found in the peat and other superficial deposits of England and the continent of Europe; while in the Pleistocene formations of England and Siberia occur remains of a giant extinct beaver, Trogontherium cuvieri, representing a genus by itself.